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aSymbols: +, characteristic positive or substrate utilized by most strains; ±, characteristic positive or substrate utilized by some strains; -not utilized by most strains; U, unknown—no data available; ( +) weak activity or growth.

bNone of the species uses formate, methanol, ethanol, propanol, citrate, or benzoate.

, characteristic negative or substrate aSymbols: +, characteristic positive or substrate utilized by most strains; ±, characteristic positive or substrate utilized by some strains; -not utilized by most strains; U, unknown—no data available; ( +) weak activity or growth.

bNone of the species uses formate, methanol, ethanol, propanol, citrate, or benzoate.

, characteristic negative or substrate

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